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Laboratoire d'Océanographie Microbienne
UMR 7621

Stéphane Blain





Pr Blain Stéphane

Professeur UPMC - HDR

Laboratoire d'Océanographie Microbienne (LOMIC) - UMR 7621 CNRS-UPMC

Avenue Fontaulé - 66650 Banyuls sur mer, France

Tél: 33 (0)4 68 88 73 44 ; Fax: 33 (0)4 68 88 73 95


research-ID : F-6917-2010


 1986-1992 Research assistant URA CNRS 322, Université de Bretagne occidentale

1993 Post-doctoral position Moss Landing Marine Laboratory CA USA

1993-1996 Research assistant URA CNRS 1513, Université de Bretagne Occidentale.

1996-2003 Associate professor UMR CNRS 6539 Université de Bretagne Occidentale.

2003-2007 : Professor Université de Marseille II (COM)

2008 : Professor Université Pierre et Marie Curie.



 Development of new analytical method for trace metal determination in sea water.

Distribution of trace metal in sea water, and interaction with phyto- and bacterioplankton.

Development of new in-situ chemical analysers and deployement at sea.



Biogeochemistry of carbon, nutrient and trace metal in the ocean.

Marine analytical chemistry.


Commitees and professional member ship

 International :

PI of “programme international de coopération scientifique »  of CNRS with Australia 2007-2009.

PI of French Australian science technology program (FAST) 2005-2007.  

Co-convener of a special session at the Ocean Science Meeting 2006.

SCOR expert working group #109 on « biogeochemistry of iron in seawater » 1999-2000

Scientific committee (chairman) of the international symposium marine analytical chemistry for monitoring and oceanographic research  1997.

Organising committee for the international symposium on hydrodynamic processes in the Southern Ocean, Brest, France, 1995.

Organizing committee for international symposium on the Circulation and biogeochemistry of water masses in the southern ocean, Brest, France, 1990.


Expertise for the FP7 of the European Commission (2008).


Serve as referee for 16 professional journals and 3 governmental funding agencies


National :

President of "Commission specialisé Ocean Atmosphère" of INSU 2014-present

Member of “Commission Specialist Ocean Atmosphere” of INSU 2002-2013

Member of “Commission National Flotte et Engin, evaluation” 2008-present

Member of national working group “renouvellement de la flotte” of Comité stratégique et technique de la flotte (CSTF). 2008-2014


National working group « new instrumentation and nutrient analysis »of the CNRS,1993.


Expertise for national Agency : CBRS, IFREMER, IRD, AERES. ANR

Expertise for private companies : AXA

Member of jury for 47 PhD defence and 6 HDR defence.



Leader of the team : “flux de matière, domaines océaniques” of UMR CNRS 6539. 1998-1999.

Leader of the team :  “ océanographie et cycles biogéochimques” of UMR CNRS 6539 2000-2001.

Leader of the “ cycles biogéochimiques” of UMR 7621 Banyuls sur mer 2008-present.

Scientific head of the station d’observation côtière de l’institut Universitaire européen de la mer (Brest): 1997-2003.

Scientfic head of the service d’observation du laboratoire Arago (Banyuls sur mer) 2008-present .

Director of the laboratory of microbial oceanography (UMR 7621 LOMIC) 2010-2013.


Field campaign


July 1993 : R/V Point Sur , ironex0, Monterey bay, USA.

September-October 1994 : N/O Atalante, FLUPAC, Equatorail Pacifique.

October-November 1995 : N/O Marion Dufresne, ANTARES 3, Southern Ocean.

Janary-February 1999 : N/O Marion Dufresne, campagne ANTARES 4 , Southern Ocean.

October-November 2000 : N/O Polarstern , campagne ANTXIII : East Atlantic Ocean.

February 2001 : N/O Atalante : POMME1, Nord East Atlantic Ocean.

October 2001 : N/O Thalassa : POMME3 , Nord East Atlantic Ocean.

March 2002 : N/O Pelagia : IRONAGES, Nord East Atlantic Ocean.

Octobre Novembre 2004 N/O Atalante campagne BIOSOPE, Pacifique sud est.

January-February 2005: N/O Marion Dufresne KEOPS Southern Ocean

January-February 2007 N/O Marion Dufresne KEOPS Southern Ocean.

August 2008 N/O Thetis LATEX0, Mediterranean Sea.

October November 2011 Marion Dufresne KEOPS2 Southern Ocean



M. Kersalé, A. Petrenko, A. M. Doglioli, F. Nencioli, J. Bouffarde,S. Blain, F. Diaz, T. Labasque, B. Queguiner, I. Dekeyser (in revision) Lateral diffusivity coefficients from the dynamics of a SF6 patch in a coastal environment. Journal of marine system



 H. Botebol, E.l Lesuisse, R. Sutak., C. Six., J.-C. Lozano, P. Schatt , V. Vergé, A. Kirilovsky, J. Morrissey, T. Léger, J.-M. Camadro., A. Gueunegues , C. Bowler, S. Blain and F.-Y. Bouget (in revision) A central role for ferritin in the day/night regulation of iron homeostasis in marine phytoplankton, PNAS.



 F. Fripiat, M. Elskens, T.W. Trull, S. Blain, A.-J. Cavagna, C. Fernandez, D. Fonseca-Batista, F. Planchon, P. Raimbault, A. Roukaert, and F. Dehairs (in revision) Significant mixed layer nitrification in a natural iron-fertilized bloom of the Southern Ocean Global Biogeochemical cycles



 A.-J. Cavagna, F. Fripiat, M. Elskens, F. Dehairs, P. Mangion, L. Chirurgien, I. Closset, M. Lasbleiz, L. Flores-Leiva, D. Cardinal, K. Leblanc, C. Fernandez, D. Lefèvre, L. Oriol, S. Blain, and B. Quéguiner ( (in revision) Biological productivity regime and associated N cycling in the vicinity of Kerguelen Island area, Southern Ocean Biogeosciences Volume: 12 Issue: 12



 F. d'Ovidio A. Della Penna, T. W. Trull, F. Nencioli, I. Pujol, M.H. Rio, Y.H. Park, C. Cotté, M. Zhou, and S. Blain (2015) The biogeochemical structuring role of horizontal stirring: Lagrangian perspectives on iron delivery downstream of the Kerguelen plateau Biogeosciences Volume: 12 Issue: 12 (accepted)



A. R. Bowie, P. van der Merwe, F. Quéroué, T. Trull, M. Fourquez, F. Planchon, G. Sarthou, F. Chever, A. T. Townsend, I. Obernosterer, J.-B. Sallée, and S. Blain (2015) Iron budgets for three distinct biogeochemical sites around the Kerguelen Archipelago (Southern Ocean) during the natural fertilisation study, KEOPS-2 Biogeosciences, 12, 4421-4445, 2015



 Marine L., S. Blain, U. Christaki, S. Monchy and I. Obernosterer (2015) Shifts in bacterial community composition associated with increased carbon cycling in a mosaic of phytoplankton bloom, the ISME journal, doi: 10.1038/ismej



 Quéroué, F.; Sarthou, G.; Planquette, H. F. Blain S. High variability in dissolved iron concentrations in the vicinity of the Kerguelen Islands (Southern Ocean) Biogeosciences Volume: 12 Issue: 12 Pages: 3869-388



 Rembauville, M.; Blain, S.; Armand, L., Quéguiner B., Salter I. (2015) Export fluxes in a naturally iron-fertilized area of the Southern Ocean - Part 2: Importance of diatom resting spores and faecal pellets for export Biogeosciences Volume: 12 Issue: 11 Pages: 3171-3195



 Rembauville, M.; Salter, I.; Leblond, N., Blain S. (2015) Export fluxes in a naturally iron-fertilized area of the Southern Ocean - Part 1: Seasonal dynamics of particulate organic carbon export from a moored sediment trap Biogeosciences Volume: 12 Issue: 11 Pages: 3153-3170



Beier S., Gálvez M.J., Molina V., Sarthou G. , Quéroué F., Blain S., Obernosterer I. 2015. The transcriptional regulation of the glyoxylate cycle in SAR11 in response to iron fertilization in the Southern Ocean. Environmental Microbiology Reports, doi: 10.1111/1758-2229.12267. 



 Blain S., Capparos J., Guéneuguès A., Obernosterer I., Oriol L. 2015. Distributions and stoichiometry of dissolved nitrogen and phosphorus in the iron-fertilized region near Kerguelen (Southern Ocean). Biogeosciences, 12, 623-635, doi:10.5194/bg-12-623-2015.



 van der Merwe P., Bowie A.R., Quéroué F., Armand L., Blain S., Chever F., Davies D., Dehairs F., Planchon F., Sarthou G., Townsend A.T., Trull T.W. 2015. Sourcing the iron in the naturally fertilised bloom around the Kerguelen Plateau: particulate trace metal dynamics. Biogeosciences, 12, 739-755, doi:10.5194/bg-12-739-2015.



 Laurenceau-Cornec E.C., Trull T.W., Davies D.M., Bray S.G., Doran J., Planchon F., Carlotti F., Jouandet M.-P., Cavagna A.-J., Waite A.M., Blain S. 2015. The relative importance of phytoplankton aggregates and zooplankton fecal pellets to carbon export: insights from free-drifting sediment trap deployments in naturally iron-fertilised waters near the Kerguelen Plateau. Biogeosciences, 12, 1007-1027, doi:10.5194/bg-12-1007-2015.



 Trull T.W., Davies D.M., Dehairs F., Cavagna A.-J., Lasbleiz M., Laurenceau-Cornec E.C., d'Ovidio F., Planchon F., Leblanc K., Quéguiner B., Blain S. 2015. Chemometric perspectives on plankton community responses to natural iron fertilisation over and downstream of the Kerguelen Plateau in the Southern Ocean. Biogeosciences, 12, 1029-1056, doi:10.5194/bg-12-1029-2015.



 Sanial V., van Beek P., Lansard B., Souhaut M., Kestenare E., d'Ovidio F., Zhou M., Blain S. 2015. Use of Ra isotopes to deduce rapid transfer of sediment-derived inputs off Kerguelen. Biogeosciences, 12, 1415-1430, doi:10.5194/bg-12-1415-2015.



 Fourquez M., Obernosterer I., Davies D.M., Trull T.W., Blain S. 2015. Microbial iron uptake in the naturally fertilized waters in the vicinity of the Kerguelen Islands: phytoplankton–bacteria interactions. Biogeosciences, 12, 1893-1906, doi:10.5194/bg-12-1893-2015.



 Obernosterer I., Fourquez M., Blain S. 2015. Fe and C co-limitation of heterotrophic bacteria in the naturally fertilized region off the Kerguelen Islands. Biogeosciences, 12, 1983-1992, doi:10.5194/bg-12-1983-2015.



 Laurenceau, E. C. Trull, T.W., Davies D. M., De La Rocha C. L., Blain. S.  2015 Phytoplankton morphology controls on aggregation processes and marine snow sinking velocities in a naturally iron-fertilized area of the Southern Ocean Marine Ecology Progress Series, 520, 35-56.



 Lasbleiz M., Leblanc K., Blain S., Ras J., Cornet-Barthaux V., Hélias Nunige S., Quéguiner B. 2014. Pigments, elemental composition (C, N, P, and Si), and stoichiometry of particulate matter in the naturally iron fertilized region of Kerguelen in the Southern Ocean. Biogeosciences, 11, 5931-5955, doi:10.5194/bg-11-5931-2014



 Jouandet M.-P., G.A. J. Jackson, F.C. Carlotti, M.P. Picheral, L.S. Stemmann, and S. Blain  (2014) Rapid formation of large aggregates during the spring bloom of Kerguelen Island: observations and model comparisons. Biogeosciences, 11, 4393-4406.



 Sanial V., van Beek P., Lansard  B., d’Ovidio F., Kestenare E., Souhaut M., Zhou M., Blain S. (2014) Study of the phytoplankton plume dynamics off Crozet Islands 2 (Southern Ocean): A geochemical-physical coupled approach. Journal of Geophysical Research-Oceans Volume: 119 Issue: 4 Pages: 2227-2237



 Lozano J.C., Schatt P., Botebol H., Vergé V., Lesuisse E., Blain S., Carré I. and Bouget F.Y. (2014) Efficient gene targeting and foreign DNA removal by homologous recombination in the picoeukaryote Ostreococcus. Plant Journal 78 Issue: 6 Pages: 1073-1083



 Fourquez, M. Blain S., Devez A. Jouenne T., Schauman A. Obernosterer I. Impacts of iron limitation on the carbon metabolism in oceanic and coastal heterotrophic bacteria. Limnol Oceanogr 59: 349-360.



 Landa, M., M. Cottrell, D. Kirchman, K. Kaiser, P.M. Medeiros, L. Tremblay, N. Batailler, J. Caparros, P. Catala, K. Escoubeyrou, L. Oriol, S. Blain and I. Obernosterer (in press) Phylogenetic and structural response of heterotrophic bacteria to dissolved organic matter of different chemical composition in a continuous culture. (2014) Environ Microbiol. 16:1668-1681.



 Blain S., Sophie R., Xiaogang X., Claustre H. and Guinet C. (2013) Instrumented elephant seals reveal the Seasonality in Chlorophyll and light-mixing regime in the iron fertilized Southern Ocean. Geophysical Research Letter, vol. 40, 1–5,



Landa, M., M. Cottrell, D. Kirchman, S. Blain and I. Obernosterer (2013). Changes in bacterial diversity in response to dissolved organic matter supply in a continuous culture experiment. Aquat Microb Ecol. 69 : 157-168. 



 Guinet, C., Xing, X., Walker, E., Monestier, p., Marchand, S., Picard, B., Jaud, T., Authier, M., Cotté, C., Dragon, A.C., Diamond, E., Antoine, D., Lovell, P., Blain, S., D’ortenzio, F., Claustre H. (2013) Calibration procedures and first data set of Southern ocean chlorophyll-a profiles collected by elephant seal equipped with a newly developed CTD-fluorescence tag. Earth System Science Data, 5, 19-29.



 Fourquez, M., Obernosterer I., Blain S. (2012) A method for the use of the radiotracer 55Fe for microautoradiography and CARD-FISH of natural bacterial communities. (2012) FEMS Microbiology Letters. 337(2), 132-139.



 Xing X, Claustre, H., Blain S., D’Ortenzio F., Antoine D., Ras J.,  Guinet, C. (2012) Quenching correction for in vivo chlorophyll fluorescence acquired by autonomous platforms: A case study with instrumented elephant seals in the Kerguelen region (Southern Ocean) Limnology and Oceanography methods 10, 2012, 483–495.



 Toulza E., Tagliabue A., Blain S., Piganeau G., (2012) Analysis of the Global Ocean Sampling (GOS) Project for Trends in Iron Uptake by Surface Ocean Microbes, PLoS ONE 7(2): e30931. doi:10.1371/journal.pone.0030931.



Bozec,Y., Merlivat,L., Baudoux,A. C., Beaumont,L., Blain,S., Bucciarelli,E., Danguy,T., Grossteffan,E., Guillot,A., Guillou,J., Repecaud,M., Treguer,P. (2011). Diurnal to inter-annual dynamics of pCO(2) recorded by a CARIOCA sensor in a temperate coastal ecosystem (2003-2009). Marine Chemistry, 126, 13-26.



 Unraveling Microbial Strategies of Iron Uptake in Marine Metagenomes. E. Toulza, A.Tagliabue, S. Blain and G. Piganeau, genomic biology submitted



 Jouandet M. P., Trull T. W., Guidi L., Picheral M., Ebersbach F., Stemmann L., Blain S. (2011). Optical imaging of mesopelagic particles indicates deep carbon flux beneath a natural iron-fertilized bloom in the Southern Ocean. Limnology and Oceanography, 56, 3, 1130-1140.



 Jouandet M. P., Blain S., Metzl N., Mongin (2011) Interannual variability of net community production and air-sea CO2 flux in a naturally iron fertilized region of the Southern Ocean (Kerguelen plateau). Antarctic Science doi:10.1017/S0954102011000411



Goberville, E., Beaugrand G., Sautour B., Tréguer, P. and SOMLIT team, Climate driven changes in coastal marine systems of western europe. Marine Ecology progress series, 408, 129-147,2010.


 Laghdass, M., S. Blain, M. Besseling, C. Guieu and I. Obernosterer: "Effects of Saharan dust on the micrbial community during a large clean mesocosm experiment in the NW Mediterranean Sea" Aquat Microb Ecol (doi10.3354/ame01466)



C. Guieu, F. Dulac, K. Desboeufs, T. Wagener, E. Pulido-Villena, J.-M. Grisoni, F. Louis, C. Ridame, S. Blain, C. Brunet, E. Bon Nguyen, S. Tran, M. Labiadh, and J.-M. Dominici.  2010. Large clean mesocosms and simulated dust deposition: a new methodology to investigate responses of marine oligotrophic ecosystems to atmospheric inputs.Biogeosciences, 7, 2765-2784, .



Chever F, Sarthou G, Bucciarelli E, Blain S, Bowie A. R. 2009. An iron budget during the natural iron fertilisation experiment KEOPS (Kerguelen Islands, Southern Ocean), Biogeosciences 6:6803-6837

 Nouvelle fenêtre


Blain, S., Quéguiner, B. and Trull, T., 2008a. The natural iron fertilization experiment KEOPS (KErguelen Ocean and Plateau compared Study): An overview. Deep Sea Research Part II: Topical Studies in Oceanography, 55(5-7): 559.


pdfNouvelle fenêtre


Blain, S., Sarthou, G. and Laan, P., 2008b. Distribution of dissolved iron during the natural iron-fertilization experiment KEOPS (Kerguelen Plateau, Southern Ocean). Deep Sea Research Part II: Topical Studies in Oceanography, 55(5-7): 594.



Bopp, L., Aumont, O., Belviso, S. and Blain, S., 2008. Modelling the effect of iron fertilization on dimethylsulphide emissions in the Southern Ocean. Deep Sea Research Part II: Topical Studies in Oceanography, 55(5-7): 901.


Gerringa, L.J.A., Blain, S., Laan, P., Sarthou, G., Veldhuis, M.J.W., Brussaard, C.P.D., Viollier, E. and Timmermans, K.R., 2008. Fe-binding dissolved organic ligands near the Kerguelen Archipelago in the Southern Ocean (Indian sector). Deep Sea Research Part II: Topical Studies in Oceanography, 55(5-7): 606.



  Jouandet, M.P., Blain, S., Metzl, N., Brunet, C., Trull, T.W. and Obernosterer, I., 2008. A seasonal carbon budget for a naturally iron-fertilized bloom over the Kerguelen Plateau in the Southern Ocean. Deep Sea Research Part II: Topical Studies in Oceanography, 55(5-7): 856.



Blain, S., Bonnet, S. and Guieu C. 2008. Dissolved iron distribution in the Southern eastern tropical and sub tropical Pacific. Biogeosciences, 5, 269-280.

pdfNouvelle fenêtre


Blain, S., B. Quéguiner, L. Armand, S. Belviso, B. Bombled, L. Bopp, A. Bowie, C. Brunet, K. Brussaard, F. Carlotti, U. Christaki, A. Corbière, I. Durand, F. Ebersbach, J.L. Fuda, N. Garcia, L.J.A. Gerringa, F.B. Griffiths, C. Guigue, C. Guillerm, S. Jacquet, C. Jeandel, P. Laan, D. Lefèvre, C. Lomonaco, A. Malits, J. Mosseri, I. Obernosterer, Y.H. Park, M. Picheral, P. Pondaven, T. Remenyi, V. Sandroni, G. Sarthou, N. Savoye, L. Scouarnec, M. Souhault, D. Thuillers, K.R. Timmermans, T. Trull, J. Uitz, P. Van-Beek, M.J.W. Veldhuis, D. Vincent, E. Viollier, L. Vong, and T. Wagener, Effect of natural iron fertilisation on carbon sequestration in the Southern Ocean. Nature, 2007, 1070-1074.



  Boyd, P.W., T. Jickells, C. Law, S. Blain, E.A. Boyle, K.O. Buesseler, K.H. Coale, J.J. Cullen, H.J.W. De Baar, M. Follows, M. Harvey, C. Lancelot, M. Levasseur, D.A. Pollard, R.B. Rivkin, J.L. Sarmiento, V. Schoemann, V. Smetacek, S. Takeda, A. Tsuda, D.R. Turner, and A. Watson, A synthesis of mesoscale iron-enrichment experiments 1993-2005: key findings and implications for ocean biogeochemistry. Science, 2007, 315, 612-617.



  Ussher, S., Worsfold, P.J., Achterberg, E., Laës, A., Blain, S., Laan, P., deBaar, P. Distribution and redox speciation of dissilved iron on the European continental margin. 2007 Limnology and Oceanopgraphy, 52(6), 2530-2539.



Laës, A., S. Blain, P. Laan, S. Ussher, E.P. Achterberg, P. Tréguer, and H.J.W. De Baar, Sources and transport of dissolved iron and manganese along the continental margin of the Bay of Biscay. 2007, Biogeoscience, 4, 181-194.



Vong, L., A. Laës, and S. Blain, Determination of iron-porphyrin-like complexes at nanomolar levels in seawater. 2007 Analytica Chimica Acta, 588, 237-244.



Sarthou, G., A.R. Baker, J. Kramer, P. Laan, A. Laës, S. Ussher, E.P. Achterberg, H.J.W. De Baar, K.R. Timmermans, and S. Blain, Influence of atmopsheric inputs on the iron distribution in the subtropical North-East Atlantic Ocean. Marine Chemistry, 2007, 104,186-202.




  Laës, A., R. Vuillemin, B. Leilde, G. Sarthou, C. Bournot-Marec, and S. Blain, Impact of environmental factors on in situ determination of iron in seawater by flow injection analysis. Marine Chemistry, 2005. 97: p. 347-356.



Sarthou, G., K.R. Timmermans, S. Blain, and P. Tréguer, Growth physiology and fate of diatoms in the ocean: a review. Journal of Sea Research, 2005. 53(1-2): p. 25.




  Blain, S., J. Guillou, P. Woerther, L. Delauney, O. Gontier, M. Hamon, B. Leildé, A. Masson, P. Tréguer, C. Tartu, and R. Vuillemin, High frequency monitoring of coastal marine environment using MAREL bouy. Journal of environmental monitoring, 2004. 6: p. 569-575.



Blain, S., C. Guieu, H. Claustre, K. Leblanc, T. Moutin, B. Quéguiner, and G. Sarthou, Availability of iron and major nutrients for phytoplankton in the north-east Atlantic ocean. Limnology and Oceanography, 2004. 49(6): p. 2095-2104. 



  Bowie, A.R., E.P. Achterberg, S. Blain, M. Boye, P.L. Croot, H.J.W. de Baar, P. Laan, G. Sarthou, and P.J. Worsfold, Shipboard analytical intercomparison of dissolved iron in surface waters along a north-south transect of the Atlantic Ocean. Marine Chemistry, 2003. 84(1-2): p. 19.



  Aumont, O., E. Maier-Reimer, S. Blain, and P. Monfray, An ecosystem model of the global ocean including Fe, Si, P co limitations. Global Biogeochemical Cycles, 2003. 17(2): p. 1060.



  Laes, A., S. Blain, P. Laan, E.P. Achterberg, G. Sarthou, and H.J.W. De Baar, Deep dissolved iron profiles in the eastern North Atlantic in relation to water masses. Geophysical Research Letters, 2003. 30(17).



  Sarthou, G., A.R. Baker, S. Blain, E.P. Achterberg, M. Boye, A.R. Bowie, P. Croot, P. Laan, H.J.W. de Baar, T.D. Jickells, and P.J. Worsfold, Atmospheric iron deposition and sea-surface dissolved iron concentrations in the eastern Atlantic Ocean. Deep Sea Research Part I: Oceanographic Research Papers, 2003. 50(10-11): p. 1339.



Sedwick, P.N., S. Blain, B. Queguiner, F.B. Griffiths, M. Fiala, E. Bucciarelli, and M. Denis, Resource limitation of phytoplankton growth in the Crozet Basin, Subantarctic Southern Ocean. Deep Sea Research Part II: Topical Studies in Oceanography, 2002. 49(16): p. 3327.



Blain, S., P.N. Sedwick, F.B. Griffiths, B. Queguiner, E. Bucciarelli, M. Fiala, P. Pondaven, and P. Treguer, Quantification of algal iron requirements in the Subantarctic Southern Ocean (Indian sector). Deep Sea Research Part II: Topical Studies in Oceanography, 2002. 49(16): p. 3255.



  Leblanc, K., B. Queguiner, M. Fiala, S. Blain, J. Morvan, and R. Corvaisier, Particulate biogenic silica and carbon production rates and particulate matter distribution in the Indian sector of the Subantarctic Ocean. Deep Sea Research Part II: Topical Studies in Oceanography, 2002. 49(16): p. 3189.



Guieu, C., Y. Bozec, S. Blain, C. Ridame, G. Sarthou, and N. Leblond, Impact of high saharan dust inputs on dissolved iron distributions in the Mediterranean sea. Geophysical Research Letters, 2002. 29(19): p. 1911.



Blain S., P. Tréguer, S.Belviso,  E. Bucciarelli, M. Denis, S. Desabre, M. Fiala , J. Le Fèvre,, V. Martin Jézéquel, J. C. Marty, P. Mayzaud, S. Razouls. (2001) Biogeochemical study of an island mass effect in the context of the iron hypothesis in the Southern Ocean. Deep Sea Research I, 48, 163-187.



Bucciarelli, E., S. Blain and P. Tréguer (2001) Distributions of iron and manganese in the wake of the Kerguelen islands, southern ocean. Marine Chemistry, 73, 21-36.



  Dunne J., J. W. Murray, A. Aufdenkampe, S. Blain and M. Rodier. (1999) Silicon-nitrogen coupling in the equatorial Pacific upwelling. Global biogeochemical cycles, 13(3), 715.



Blain S.,  Tréguer P., Rodier M., (1999). Stocks and fluxes of biogenic silica in the western oligotrophic equatorial Pacific. Journal of Geophysical Research, 104 C2, 3357-3367.



  Floch J., Blain S., Birot D., Tréguer P., (1998). In-situ determination of silicic acid in seawater based on FIA and colorimetric dual-wavelength measurements. Analytica Chimica Acta, 377, 157-166.



Coale K.H., Johnson K.S., Fitzwater S.E., Blain S., Stanton T.P., Coley T.L., (1998). IronEx-I, an in-situ iron-enrichment experiment : experimental design, implementation and results. Deep-Sea research II, 45, 919-945.



Blain, S., A. Leynaert, P. Tréguer, M.J. Chrétiennot Dinet and M. Rodier. (1997) Biomass, growth rates and limitation of equatorial Pacific diatoms. Deep Sea research, 44 (7), 1255-1275.



Daniel, A., D. Birot, S. Blain, P. Tréguer, E. Menut and B. Leilde. (1995) A submersible flow-injection analyser for the in-situ determination of nitrite and nitrate in coastal waters. Marine Chemistry 51, 67-77.



Blain, S., Tréguer P. (1995) Iron(II) and iron(III) determination in seawater at the nanomolar level with on line preconcentration and spectrophotometric detection. Analytica Chimica Acta, 308, 425-432.



  Blain, S., P. Appriou and H. Handel. (1993) Preconcentration of trace metal from seawater with a chelating resin: chelamine. Analytica Chimica Acta. 272, 91-97.



Blain, S., P Appriou and H. Handel(1991). Column preconcentration of trace metals from seawater with macroporous resins impregnated with lipophilic tetraazamacrocycles. Analyst, 116, 815-820.



  Blain, S., H. Chaumeil, P. Appriou and H. Handel (1990). Application of a tetrazamacrocycle immobilized on an organic polymer to the determination of trace amounts of manganese in seawater. Analytica chimica Acta. 232, 331-336.


Traductions :

    Le LOMIC en chiffres

    6 Chercheurs CNRS
    4 Enseignant-chercheurs
    8 Personnels techniques
    4 Post doctorants
    8 Etudiants en thèse

    4 Etudiants Master2