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Laboratoire d'Océanographie Microbienne
UMR 7621

Stéphane BLAIN



Pr BLAIN Stéphane


Professeur Sorbonne Université - HDR


Laboratoire d'Océanographie Microbienne (LOMIC) - UMR 7621 CNRS-SU


Avenue Fontaulé - 66650 Banyuls sur mer, France


Tél: 33 (0)4 68 88 73 44


E-mail: stephane.blain @


Research-ID : F-6917-2010


Main Achievements

Stéphane Blain received his Ph.D. in chemical oceanography from the University of Brest in 1992. In 1993, he served as a post doc at the Moss Landing Marine Laboratories during the preparation of IRONEX1. He is actually professor of chemical oceanography at the University Pierre et Marie Curie (Paris). He is working at the Marine station of Banyuls sur Mer, where he has founded and directed the Microbial Oceanography Laboratory (2010-2013), a research unit in partnership with the UPMC and the CNRS. His research interests are the functioning of the biological pump of CO2 in the ocean with focus on iron biogeochemistry and the Southern Ocean. He is also involved in the development of new analytical tools and methods for in-situ chemical oceanography.

He has initiated the project KEOPS (KEOPS: Kerguelen Ocean and Plateau compared study) ten years ago. The main objective of KEOPS is to study the impact of natural iron fertilization of the Southern Ocean. KEOPS is international project that gathered researchers from 10 French laboratories and foreign colleagues from different nations (Australia, Belgium, Chili, The Netherlands, South Korea, USA, UK). The first phase of this project was conducted in 2004-2009 and and the second phase took place in 2011-2014). All together more than 100 publication were produced.. As principal investigator of KEOPS, He has co-organized two special sessions with colleagues of the National Oceanographic Center of Southampton (OSM, Hawaii 2006) and with colleagues of the Alfred Wegener Institute (ASLO meeting, Orlando 2013). He was involved in the SCOR WG and invited to international workshops and conferences. The strong collaboration with australien colleagues during KEOPS has led to the invitation of S. Blain as invited professor of Tasmania University in 2015 and to the participation at the HEOBI cruise (Jan-Feb 2016) at Heard Island.

During the past 20 years he has been very interested in observing systems. In 1997, he has organized an international symposium on marine analytical chemistry for the monitoring of oceanographic research. He has participated to the first development of nutrient in situ chemical analyzers and to the deployments of biogeochemical sensors on buoys. Recently, he has used fluorescence sensors mounted on elephant seals of Kerguelen to produce the first two dimensional climatology of chlorophyll in a region of the Southern Ocean. He is presently Co-PI of the project SOCLIM (Southern Ocean and Climate) dedicated to the study of the carbon cycle in the Southern, using innovative tools like BGC-Argo floats and instrumented moorings.

Besides the research activities described above S. Blain has also explored the multiple facets of the interactions between iron and microorganisms using laboratory experiments. He was the PI of the ANR BACCIO (Biomolecular approach of the iron and carbon cycle in the ocean) project where the metabolic response of heterotrophic bacteria to iron limiation was extensively studied. In collaboration with other colleagues of the laboratory he has for the first time applied genetic tools (mutants and bioreporters) to elucidate the role of ferrritine in iron homeostasis of picophytoplancton. New studies are now underway to apply the same approaches to heterotrophic bacteria.



2001 : Habilitation à diriger des recherches (french degree mandatory for supervising PhD)

1992 :PhD in marine analytical chemistry Université de Bretagne Occidentale, Brest, France

1988: DEA (now called master) Université de Bretagne Occidentale, Brest, France

1986 agrégation of chimie (french diploma of highest level for full-time teaching).

1983 CAPES of Physic and chemistry (French diploma for full-time teaching in high school).



2008 : Professor Université Pierre et Marie Curie.

2003-2007 : Professor Université de Marseille II (COM)

1996-2003 Associate professor UMR CNRS 6539 Université de Bretagne Occidentale.

1993-1996 Research assistant URA CNRS 1513, Université de Bretagne Occidentale.

1993 Post-doctoral position Moss Landing Marine Laboratory CA USA

1986-1992 Research assistant URA CNRS 322, Université de Bretagne occidentale


Recent award

Invited professor at University of Tasmania and at the Institut for antarctic studies (Hobart, Australia)


Field experience

July 1993 : R/V Point Sur , ironex0, Monterey bay, USA.

September-October 1994 : N/O Atalante, FLUPAC, Equatorail Pacifique.

October-November 1995 : N/O Marion Dufresne, ANTARES 3, Southern Ocean.

Janary-February 1999 : N/O Marion Dufresne, campagne ANTARES 4 , Southern Ocean.

October-November 2000 : N/O Polarstern , campagne ANTXIII : East Atlantic Ocean.

February 2001 : N/O Atalante : POMME1, Nord East Atlantic Ocean.

October 2001 : N/O Thalassa : POMME3 , Nord East Atlantic Ocean.

March 2002 : N/O Pelagia : IRONAGES, Nord East Atlantic Ocean.

Octobre Novembre 2004 N/O Atalante campagne BIOSOPE, Pacifique sud est.

January-February 2005: N/O Marion Dufresne KEOPS Southern Ocean

January-February 2007 N/O Marion Dufresne KEOPS Southern Ocean.

August 2008 N/O Thetis LATEX0, Mediterranean Sea.

October November 2011 Marion Dufresne KEOPS2 Southern Ocean

January-February 2015 Investigator HEOBI, Southern Ocean

October 2016 Marion Dufresne SOCLIM Southern Ocean

March 2017 Marion Dufresne SOCLIM Southern Ocean

February March 2018 (planed) Marion Dufresne MOBYDICK Southern Ocean


Chair and Membership

International :

PI of “programme international de coopération scientifique »  of CNRS with Australia 2007-2009.

PI of French Australian science technology program (FAST) 2005-2007.  

Co-convener of a special session at the Ocean Science Meeting 2006.

SCOR expert working group #109 on « biogeochemistry of iron in seawater » 1999-2000

Scientific committee (chairman) of the international symposium marine analytical chemistry for monitoring and oceanographic research  1997.

Organising committee for the international symposium on hydrodynamic processes in the Southern Ocean, Brest, France, 1995.

Organizing committee for international symposium on the Circulation and biogeochemistry of water masses in the southern ocean, Brest, France, 1990. 

Expertise for the FP7 of the European Commission (2008). 

Serve as referee for 16 professional journals and 3 governmental funding agencies 


National :

President of "Commission specialisée Ocean Atmosphère" of INSU 2014-2017

Member of “Commission Specialisée Ocean Atmosphere” of INSU 2002-2013

Member of “Commission National Flotte et Engin, evaluation” 2008-present

Member of national working group “renouvellement de la flotte” of Comité stratégique et technique de la flotte (CSTF). 2008-2014 

National working group « new instrumentation and nutrient analysis »of the CNRS,1993. 

Expertise for national Agency : CBRS, IFREMER, IRD, AERES. ANR

Expertise for private companies : AXA

Member of jury for 47 PhD defence and 6 HDR defence. 


Local :

Leader of the team : “flux de matière, domaines océaniques” of UMR CNRS 6539. 1998-1999.

Leader of the team :  “ océanographie et cycles biogéochimques” of UMR CNRS 6539 2000-2001.

Leader of the “ cycles biogéochimiques” of UMR 7621 Banyuls sur mer 2008-present.

Scientific head of the station d’observation côtière de l’institut Universitaire européen de la mer (Brest): 1997-2003.

Scientific head of the service d’observation du laboratoire Arago (Banyuls sur mer) 2008-2010 .

Director of the laboratory of microbial oceanography (UMR 7621 LOMIC) 2010-2013.


Supervision of graduate students and postdoctoral fellows.

Supervision of 12 PhD, 6 post doc, 15 Master.


            1993-1994 Daniel, A., PhD University of Brest. co-supervisor.

            1994-1998 Floch, J., University of Brest  supervisor.

            1998-2001 Bucciarelli, E. , University of Brest supervisor

            2001-2003 Laes A., University of Brest supervisor

            2004-2007 Vong L.  University of Marseille supervisor

            2005-2008 Wagener , T., University of Paris 6 co-supervisor

            2005-2008 Jouandet M.-P., University of Marseille, co-supervisor

            2009-2012 Fourquez M. University of Paris 6 supervisor

            2010-2013 Landa M. University of Paris 6 co supervisor

            2011-2014 Botebol H. University of Paris 6 co supervisor

            2012-2016 Rembauville M.  University of Paris 6 co supervisor

            2016-2018 Koedooder C. University of Paris supervisor

Post doctoral fellows

            2003 Sarthou, G., University of Brest France (project IRONAGES FP5)

            2007-2008 Xing, X., (project IPOS-SEAL ANR)

            2009 Devez A. (project BACCIO ANR)

            2009 Toulza E. (project BACCIO ANR)

            2010-2012 Beier, S. (projects BACCIO and KEOPS2)

            2016 Rembauville (project SOCLIM)


Teaching activities

At the University of Brest, I have taught general chemistry, Thermodynamics and quantum chemistry, aquatic chemistry, analytical chemistry.

At the University of Marseille, I have taught analytical chemistry, chemical oceanography, chemical sensors for oceanography, carbon cycle in the ocean, geochemical tracers.

At the University of Paris 6, I teach marine observing systems, marine chemistry, biogeochemistry and microbes in the ocean, the practice of Ocean Data View.

At the 3 universities, I have organized practical courses based on short cruises on research vessels for collecting water samples in coastal environments. The samples were then analyzed during practical courses. Finally the results were discussed and presented by the students.


Broad audience dissiminetation.

I have contributed to disseminate my findings in the media with 15 articles and interviews in scientific magazines, 10 interviews in national newspapers, participations to 9 radio and to 2 television emissions,  participation to the movie “ CO2 public enemy N°1” by Nicolas Koutiskas in 2009.

The most recent achievement in this field is the web TV series SOCLIM


Peer Review pubilcations


Rembauville, M., Briggs, N., Ardyna, M., Uitz, J., Catala, P., Penkerc’ch, C., Poteau, A., Claustre, H., and Blain, S. (2017) Plankton assemblage estimated with BGC-Argo floats in the Southern Ocean: implications for seasonal successions and particle export in the mesopelagic. Journal of Geophysical Research : Oceans. doi:10.1002/2017JC013067


Rembauville, M., Salter, I., Dehairs, F. , Miquel, J.-C. and Blain, S.:(2017) Annual particulate matter and diatom export in a high nutrient, low chlorophyll area of the Southern Ocean. Polar Biology. doi:10.1007/s00300-017-2167-3


Botebol, H., Lelandais, G., Six, C., Lesuisse, E., Meng, A., Bittner, L., Blain S., Bouget, F.-Y. (2017). Acclimation of a low iron adapted Ostreococcus strain to iron limitation through cell biomass lowering. Scientific Reports, 7(1), 327.


Rembauville, M.; Meilland, J.; Ziveri, R., Schiebel R., Blain S. , Salter I. Planktic foraminifer and coccolith contribution to carbonate export fluxes over the central Kerguelen Plateau (2016) Deep-Sea Research Part I-Oceanographic Research Papers ,  91-101


Rembauville, M.; Manno, C.; Tarling, G. A, Blain S., Salter I.Strong contribution of diatom resting spores to deep-sea carbon transfer in naturally iron-fertilized waters downstream of South Georgia(2016)  Deep-Sea Research Part I-Oceanographic Research Papers 115 Pages: 22-35


Closset, I.; Cardinal, D.; Rembauville, M.; Thil, S., Blain S. Unveiling the Si cycle using isotopes in an iron-fertilized zone of the Southern Ocean: from mixed-layer supply to export (2016)Biogeosceinces: 13 , 21 Pages: 6049-6066


Rembauville, M.; Blain, S.; Caparros, J.; Salter I..Particulate matter stoichiometry driven by microplankton community structure in summer in the Indian sector of the Southern Ocean(2016) Limnology and Oceanography Volume: 61 Issue: 4 Pages: 1301-1321


 M. Kersalé, A. Petrenko, A. M. Doglioli, F. Nencioli, J. Bouffarde,,S. Blain, F. Diaz, T. Labasque, B. Queguiner, I. Dekeyser (2016) Lateral diffusivity coefficients from the dynamics of a SF6 patch in a coastal environment. Journal of Marine Systems Volume: 153 Pages: 42-54


 H. Botebol, E.l Lesuisse, R. Sutak., C. Six., J.-C. Lozano, P. Schatt , V. Vergé, A. Kirilovsky, J. Morrissey, T. Léger, J.-M. Camadro., A. Gueunegues , C. Bowler, S. Blain and F.-Y. Bouget (2015) A central role for ferritin in the day/night regulation of iron homeostasis in marine phytoplankton, PNAS 112,47.


 F. Fripiat, M. Elskens, T.W. Trull, S. Blain, A.-J. Cavagna, C. Fernandez, D. Fonseca-BatistaF. Planchon, P. Raimbault, A. Roukaert, and F. Dehairs 2015) Significant mixed layer nitrification in a natural iron-fertilized bloom of the Southern Ocean Global Biogeochemical Cycles Volume: 29 Issue: 11 Pages: 1929-1943


 A.-J. Cavagna, F. Fripiat, M. Elskens, F. Dehairs, P. Mangion, L. Chirurgien, I. Closset, M. Lasbleiz, L. Flores-Leiva, D. Cardinal, K. Leblanc, C. Fernandez, D. Lefèvre (2015), L. Oriol, S. Blain, and B. Quéguiner ( (in revision) Biological productivity regime and associated N cycling in the vicinity of Kerguelen Island area, Southern Ocean Biogeosciences Volume: 12 Issue: 12


 F. d'Ovidio A. Della Penna, T. W. Trull, F. Nencioli, I. Pujol, M.H. Rio, Y.H. Park, C. Cotté, M. Zhou, andS. Blain (2015) The biogeochemical structuring role of horizontal stirring: Lagrangian perspectives on iron delivery downstream of the Kerguelen plateau Biogeosciences Volume: 12 Issue: 12 (accepted)


A. R. Bowie, P. van der Merwe, F. Quéroué, T. Trull, M. Fourquez, F. Planchon, G. Sarthou, F. Chever, A. T. Townsend, I. Obernosterer, J.-B. Sallée, and S. Blain (2015) Iron budgets for three distinct biogeochemical sites around the Kerguelen Archipelago (Southern Ocean) during the natural fertilisation study, KEOPS-2 Biogeosciences, 12, 4421-4445, 2015


 Landa M.,  S. Blain, U. Christaki, S. Monchy and I. Obernosterer (2015) Shifts in bacterial community composition associated with increased carbon cycling in a mosaic of phytoplankton bloom, the ISME journal Isme Journal Volume: 10 Issue: 1 Pages: 39-50


 Quéroué, F.; Sarthou, G.; Planquette, H. F. Blain S. (2015) High variability in dissolved iron concentrations in the vicinity of the Kerguelen Islands (Southern Ocean) Biogeosciences Volume: 12 Issue: 12 Pages: 3869-388


 Rembauville, M.; Blain, S.; Armand, L., Quéguiner B., Salter I. (2015) Export fluxes in a naturally iron-fertilized area of the Southern Ocean - Part 2: Importance of diatom resting spores and faecal pellets for export Biogeosciences Volume: 12 Issue: 11 Pages: 3171-3195


 Rembauville, M.; Salter, I.; Leblond, N., Blain S. (2015) Export fluxes in a naturally iron-fertilized area of the Southern Ocean - Part 1: Seasonal dynamics of particulate organic carbon export from a moored sediment trap Biogeosciences Volume: 12 Issue: 11 Pages: 3153-3170


Beier S., Gálvez M.J., Molina V., Sarthou G. , Quéroué F., Blain S., Obernosterer I. 2015. The transcriptional regulation of the glyoxylate cycle in SAR11 in response to iron fertilization in the Southern Ocean. Environmental Microbiology Reports, doi: 10.1111/1758-2229.12267. 


 Blain S., Capparos J., Guéneuguès A., Obernosterer I., Oriol L. 2015. Distributions and stoichiometry of dissolved nitrogen and phosphorus in the iron-fertilized region near Kerguelen (Southern Ocean). Biogeosciences12, 623-635, doi:10.5194/bg-12-623-2015.


 van der Merwe P., Bowie A.R., Quéroué F., Armand L., Blain S., Chever F., Davies D., Dehairs F., Planchon F., Sarthou G., Townsend A.T., Trull T.W. 2015. Sourcing the iron in the naturally fertilised bloom around the Kerguelen Plateau: particulate trace metal dynamics. Biogeosciences12, 739-755, doi:10.5194/bg-12-739-2015.


 Laurenceau-Cornec E.C., Trull T.W., Davies D.M., Bray S.G., Doran J., Planchon F., Carlotti F., Jouandet M.-P., Cavagna A.-J., Waite A.M., Blain S. 2015. The relative importance of phytoplankton aggregates and zooplankton fecal pellets to carbon export: insights from free-drifting sediment trap deployments in naturally iron-fertilised waters near the Kerguelen Plateau. Biogeosciences12, 1007-1027, doi:10.5194/bg-12-1007-2015.


 Trull T.W., Davies D.M., Dehairs F., Cavagna A.-J., Lasbleiz M., Laurenceau-Cornec E.C., d'Ovidio F., Planchon F., Leblanc K., Quéguiner B., Blain S. 2015. Chemometric perspectives on plankton community responses to natural iron fertilisation over and downstream of the Kerguelen Plateau in the Southern Ocean. Biogeosciences, 12, 1029-1056, doi:10.5194/bg-12-1029-2015.


 Sanial V., van Beek P., Lansard B., Souhaut M., Kestenare E., d'Ovidio F., Zhou M., Blain S. 2015. Use of Ra isotopes to deduce rapid transfer of sediment-derived inputs off Kerguelen. Biogeosciences12, 1415-1430, doi:10.5194/bg-12-1415-2015.


 Fourquez M., Obernosterer I., Davies D.M., Trull T.W., Blain S. 2015. Microbial iron uptake in the naturally fertilized waters in the vicinity of the Kerguelen Islands: phytoplankton–bacteria interactions.Biogeosciences12, 1893-1906, doi:10.5194/bg-12-1893-2015.


 Obernosterer I., Fourquez M., Blain S. 2015. Fe and C co-limitation of heterotrophic bacteria in the naturally fertilized region off the Kerguelen Islands. Biogeosciences, 12, 1983-1992, doi:10.5194/bg-12-1983-2015.


 Laurenceau, E. C. Trull, T.W., Davies D. M., De La Rocha C. L., Blain. S.  2015 Phytoplankton morphology controls on aggregation processes and marine snow sinking velocities in a naturally iron-fertilized area of the Southern Ocean Marine Ecology Progress Series, 520, 35-56.


 Lasbleiz M., Leblanc K., Blain S., Ras J., Cornet-Barthaux V., Hélias Nunige S., Quéguiner B. 2014. Pigments, elemental composition (C, N, P, and Si), and stoichiometry of particulate matter in the naturally iron fertilized region of Kerguelen in the Southern Ocean. Biogeosciences11, 5931-5955, doi:10.5194/bg-11-5931-2014


 Jouandet M.-P., G.A. J. Jackson, F.C. Carlotti, M.P. Picheral, L.S. Stemmann, and S. Blain  (2014)Rapid formation of large aggregates during the spring bloom of Kerguelen Island: observations and model comparisons. Biogeosciences, 11, 4393-4406.


 Sanial V., van Beek P., Lansard  B., d’Ovidio F., Kestenare E., Souhaut M., Zhou M., Blain S. (2014)Study of the phytoplankton plume dynamics off Crozet Islands 2 (Southern Ocean): A geochemical-physical coupled approach. Journal of Geophysical Research-Oceans Volume: 119 Issue: 4 Pages: 2227-2237


 Lozano J.C., Schatt P., Botebol H., Vergé V., Lesuisse E., Blain S., Carré I. and Bouget F.Y. (2014)Efficient gene targeting and foreign DNA removal by homologous recombination in the picoeukaryote Ostreococcus. Plant Journal 78 Issue: 6 Pages: 1073-1083


 Fourquez, M. Blain S., Devez A. Jouenne T., Schauman A. Obernosterer I. (2014) Impacts of iron limitation on the carbon metabolism in oceanic and coastal heterotrophic bacteria. Limnol Oceanogr 59: 349-360.


 Landa, M., M. Cottrell, D. Kirchman, K. Kaiser, P.M. Medeiros, L. Tremblay, N. Batailler, J. Caparros, P. Catala, K. Escoubeyrou, L. Oriol, S. Blain and I. Obernosterer (in press) Phylogenetic and structural response of heterotrophic bacteria to dissolved organic matter of different chemical composition in a continuous culture. (2014) Environ Microbiol. 16:1668-1681.


 Blain S., Sophie R., Xiaogang X., Claustre H. and Guinet C. (2013) Instrumented elephant seals revealthe Seasonality in Chlorophyll and light-mixing regime in the iron fertilized Southern Ocean. Geophysical Research Letter, vol. 40, 1–5,


Landa, M., M. Cottrell, D. Kirchman, S. Blain and I. Obernosterer (2013). Changes in bacterial diversity in response to dissolved organic matter supply in a continuous culture experiment. Aquat Microb Ecol. 69 : 157-168. 


 Guinet, C., Xing, X., Walker, E., Monestier, p., Marchand, S., Picard, B., Jaud, T., Authier, M., Cotté, C., Dragon, A.C., Diamond, E., Antoine, D., Lovell, P., Blain, S., D’ortenzio, F., Claustre H. (2013)Calibration procedures and first data set of Southern ocean chlorophyll-a profiles collected by elephant seal equipped with a newly developed CTD-fluorescence tag. Earth System Science Data, 5, 19-29.


 Fourquez, M., Obernosterer I., Blain S. (2012) A method for the use of the radiotracer 55Fe for microautoradiography and CARD-FISH of natural bacterial communities. (2012) FEMS Microbiology Letters. 337(2), 132-139.


 Xing X, Claustre, H., Blain S., D’Ortenzio F., Antoine D., Ras J.,  Guinet, C. (2012) Quenching correction for in vivo chlorophyll fluorescence acquired by autonomous platforms: A case study with instrumented elephant seals in the Kerguelen region (Southern Ocean) Limnology and Oceanography methods 10, 2012, 483–495.


 Toulza E., Tagliabue A., Blain S., Piganeau G., (2012) Analysis of the Global Ocean Sampling (GOS) Project for Trends in Iron Uptake by Surface Ocean Microbes, PLoS ONE 7(2): e30931. doi:10.1371/journal.pone.0030931.


Bozec,Y., Merlivat,L., Baudoux,A. C., Beaumont,L., Blain,S., Bucciarelli,E., Danguy,T., Grossteffan,E., Guillot,A., Guillou,J., Repecaud,M., Treguer,P. (2011). Diurnal to inter-annual dynamics of pCO(2) recorded by a CARIOCA sensor in a temperate coastal ecosystem (2003-2009)Marine Chemistry,126, 13-26.


 Unraveling Microbial Strategies of Iron Uptake in Marine Metagenomes. E. Toulza, A.Tagliabue, S. Blain and G. Piganeau, genomic biology submitted


 Jouandet M. P., Trull T. W., Guidi L., Picheral M., Ebersbach F., Stemmann L., Blain S. (2011). Optical imaging of mesopelagic particles indicates deep carbon flux beneath a natural iron-fertilized bloom in the Southern OceanLimnology and Oceanography, 56, 3, 1130-1140.


 Jouandet M. P., Blain S., Metzl N., Mongin (2011) Interannual variability of net community production and air-sea CO2 flux in a naturally iron fertilized region of the Southern Ocean (Kerguelen plateau).Antarctic Science doi:10.1017/S0954102011000411


Goberville, E., Beaugrand G., Sautour B., Tréguer, P. and SOMLIT team, Climate driven changes in coastal marine systems of western europe. Marine Ecology progress series, 408, 129-147,2010.


 Laghdass, M., S. Blain, M. Besseling, C. Guieu and I. Obernosterer: "Effects of Saharan dust on the micrbial community during a large clean mesocosm experiment in the NW Mediterranean Sea" Aquat Microb Ecol (doi10.3354/ame01466)


C. Guieu, F. Dulac, K. Desboeufs, T. Wagener, E. Pulido-Villena, J.-M. Grisoni, F. Louis, C. Ridame, S. Blain, C. Brunet, E. Bon Nguyen, S. Tran, M. Labiadh, and J.-M. Dominici.  2010. Large clean mesocosms and simulated dust deposition: a new methodology to investigate responses of marine oligotrophic ecosystems to atmospheric inputs.Biogeosciences, 7, 2765-2784, .


Chever F, Sarthou G, Bucciarelli E, Blain S, Bowie A. R. 2009. An iron budget during the natural iron fertilisation experiment KEOPS (Kerguelen Islands, Southern Ocean), Biogeosciences 6:6803-6837


Blain, S., Quéguiner, B. and Trull, T., 2008a. The natural iron fertilization experiment KEOPS (KErguelen Ocean and Plateau compared Study): An overview. Deep Sea Research Part II: Topical Studies in Oceanography, 55(5-7): 559.



Blain, S., Sarthou, G. and Laan, P., 2008b. Distribution of dissolved iron during the natural iron-fertilization experiment KEOPS (Kerguelen Plateau, Southern Ocean). Deep Sea Research Part II: Topical Studies in Oceanography, 55(5-7): 594.


Bopp, L., Aumont, O., Belviso, S. and Blain, S., 2008. Modelling the effect of iron fertilization on dimethylsulphide emissions in the Southern Ocean. Deep Sea Research Part II: Topical Studies in Oceanography, 55(5-7): 901.


Gerringa, L.J.A., Blain, S., Laan, P., Sarthou, G., Veldhuis, M.J.W., Brussaard, C.P.D., Viollier, E. and Timmermans, K.R., 2008. Fe-binding dissolved organic ligands near the Kerguelen Archipelago in the Southern Ocean (Indian sector). Deep Sea Research Part II: Topical Studies in Oceanography, 55(5-7): 606.


  Jouandet, M.P., Blain, S., Metzl, N., Brunet, C., Trull, T.W. and Obernosterer, I., 2008. A seasonal carbon budget for a naturally iron-fertilized bloom over the Kerguelen Plateau in the Southern Ocean. Deep Sea Research Part II: Topical Studies in Oceanography, 55(5-7): 856.


Blain, S., Bonnet, S. and Guieu C. 2008. Dissolved iron distribution in the Southern eastern tropical and sub tropical Pacific. Biogeosciences, 5, 269-280.


Blain, S., B. Quéguiner, L. Armand, S. Belviso, B. Bombled, L. Bopp, A. Bowie, C. Brunet, K. Brussaard, F. Carlotti, U. Christaki, A. Corbière, I. Durand, F. Ebersbach, J.L. Fuda, N. Garcia, L.J.A. Gerringa, F.B. Griffiths, C. Guigue, C. Guillerm, S. Jacquet, C. Jeandel, P. Laan, D. Lefèvre, C. Lomonaco, A. Malits, J. Mosseri, I. Obernosterer, Y.H. Park, M. Picheral, P. Pondaven, T. Remenyi, V. Sandroni, G. Sarthou, N. Savoye, L. Scouarnec, M. Souhault, D. Thuillers, K.R. Timmermans, T. Trull, J. Uitz, P. Van-Beek, M.J.W. Veldhuis, D. Vincent, E. Viollier, L. Vong, and T. Wagener, Effect of natural iron fertilisation on carbon sequestration in the Southern Ocean. Nature, 2007, 1070-1074.


  Boyd, P.W., T. Jickells, C. Law, S. Blain, E.A. Boyle, K.O. Buesseler, K.H. Coale, J.J. Cullen, H.J.W. De Baar, M. Follows, M. Harvey, C. Lancelot, M. Levasseur, D.A. Pollard, R.B. Rivkin, J.L. Sarmiento, V. Schoemann, V. Smetacek, S. Takeda, A. Tsuda, D.R. Turner, and A. Watson, A synthesis of mesoscale iron-enrichment experiments 1993-2005: key findings and implications for ocean biogeochemistry.Science, 2007, 315, 612-617.


  Ussher, S., Worsfold, P.J., Achterberg, E., Laës, A., Blain, S., Laan, P., deBaar, P. Distribution and redox speciation of dissilved iron on the European continental margin. 2007 Limnology and Oceanopgraphy, 52(6), 2530-2539.


Laës, A., S. Blain, P. Laan, S. Ussher, E.P. Achterberg, P. Tréguer, and H.J.W. De Baar, Sources and transport of dissolved iron and manganese along the continental margin of the Bay of Biscay. 2007, Biogeoscience, 4, 181-194.


Vong, L., A. Laës, and S. BlainDetermination of iron-porphyrin-like complexes at nanomolar levels in seawater. 2007 Analytica Chimica Acta, 588, 237-244.


Sarthou, G., A.R. Baker, J. Kramer, P. Laan, A. Laës, S. Ussher, E.P. Achterberg, H.J.W. De Baar, K.R. Timmermans, and S. BlainInfluence of atmopsheric inputs on the iron distribution in the subtropical North-East Atlantic Ocean. Marine Chemistry, 2007, 104,186-202.


  Laës, A., R. Vuillemin, B. Leilde, G. Sarthou, C. Bournot-Marec, and S. BlainImpact of environmental factors on in situ determination of iron in seawater by flow injection analysis. Marine Chemistry, 2005.97: p. 347-356.


Sarthou, G., K.R. Timmermans, S. Blain, and P. Tréguer, Growth physiology and fate of diatoms in the ocean: a review. Journal of Sea Research, 2005. 53(1-2): p. 25.



  Blain, S., J. Guillou, P. Woerther, L. Delauney, O. Gontier, M. Hamon, B. Leildé, A. Masson, P. Tréguer, C. Tartu, and R. Vuillemin, High frequency monitoring of coastal marine environment using MAREL bouy. Journal of environmental monitoring, 2004. 6: p. 569-575.


Blain, S., C. Guieu, H. Claustre, K. Leblanc, T. Moutin, B. Quéguiner, and G. Sarthou, Availability of iron and major nutrients for phytoplankton in the north-east Atlantic ocean. Limnology and Oceanography, 2004. 49(6): p. 2095-2104. 


  Bowie, A.R., E.P. Achterberg, S. Blain, M. Boye, P.L. Croot, H.J.W. de Baar, P. Laan, G. Sarthou, and P.J. Worsfold, Shipboard analytical intercomparison of dissolved iron in surface waters along a north-south transect of the Atlantic Ocean. Marine Chemistry, 2003. 84(1-2): p. 19.


  Aumont, O., E. Maier-Reimer, S. Blain, and P. Monfray, An ecosystem model of the global ocean including Fe, Si, P co limitations. Global Biogeochemical Cycles, 2003. 17(2): p. 1060.


  Laes, A., S. Blain, P. Laan, E.P. Achterberg, G. Sarthou, and H.J.W. De Baar, Deep dissolved iron profiles in the eastern North Atlantic in relation to water masses. Geophysical Research Letters, 2003.30(17).


  Sarthou, G., A.R. Baker, S. Blain, E.P. Achterberg, M. Boye, A.R. Bowie, P. Croot, P. Laan, H.J.W. de Baar, T.D. Jickells, and P.J. Worsfold, Atmospheric iron deposition and sea-surface dissolved iron concentrations in the eastern Atlantic Ocean. Deep Sea Research Part I: Oceanographic Research Papers, 2003. 50(10-11): p. 1339.


Sedwick, P.N., S. Blain, B. Queguiner, F.B. Griffiths, M. Fiala, E. Bucciarelli, and M. Denis, Resource limitation of phytoplankton growth in the Crozet Basin, Subantarctic Southern Ocean. Deep Sea Research Part II: Topical Studies in Oceanography, 2002. 49(16): p. 3327.


Blain, S., P.N. Sedwick, F.B. Griffiths, B. Queguiner, E. Bucciarelli, M. Fiala, P. Pondaven, and P. Treguer, Quantification of algal iron requirements in the Subantarctic Southern Ocean (Indian sector).Deep Sea Research Part II: Topical Studies in Oceanography, 2002. 49(16): p. 3255.


  Leblanc, K., B. Queguiner, M. Fiala, S. Blain, J. Morvan, and R. Corvaisier, Particulate biogenic silica and carbon production rates and particulate matter distribution in the Indian sector of the Subantarctic Ocean. Deep Sea Research Part II: Topical Studies in Oceanography, 2002. 49(16): p. 3189.


Guieu, C., Y. Bozec, S. Blain, C. Ridame, G. Sarthou, and N. Leblond, Impact of high saharan dust inputs on dissolved iron distributions in the Mediterranean sea. Geophysical Research Letters, 2002.29(19): p. 1911.


Blain S., P. Tréguer, S.Belviso,  E. Bucciarelli, M. Denis, S. Desabre, M. Fiala , J. Le Fèvre,, V. Martin Jézéquel, J. C. Marty, P. Mayzaud, S. Razouls. (2001) Biogeochemical study of an island mass effect in the context of the iron hypothesis in the Southern Ocean. Deep Sea Research I, 48, 163-187.


Bucciarelli, E., S. Blain and P. Tréguer (2001) Distributions of iron and manganese in the wake of the Kerguelen islands, southern ocean. Marine Chemistry, 73, 21-36.


  Dunne J., J. W. Murray, A. Aufdenkampe, S. Blain and M. Rodier. (1999) Silicon-nitrogen coupling in the equatorial Pacific upwelling. Global biogeochemical cycles13(3), 715.


Blain S.,  Tréguer P., Rodier M., (1999). Stocks and fluxes of biogenic silica in the western oligotrophic equatorial Pacific. Journal of Geophysical Research104 C2, 3357-3367.


  Floch J., Blain S., Birot D., Tréguer P., (1998). In-situ determination of silicic acid in seawater based on FIA and colorimetric dual-wavelength measurements. Analytica Chimica Acta377, 157-166.


Coale K.H., Johnson K.S., Fitzwater S.E., Blain S., Stanton T.P., Coley T.L., (1998). IronEx-I, an in-situiron-enrichment experiment : experimental design, implementation and results. Deep-Sea research II,45, 919-945.


Blain, S., A. Leynaert, P. Tréguer, M.J. Chrétiennot Dinet and M. Rodier. (1997) Biomass, growth rates and limitation of equatorial Pacific diatoms. Deep Sea research, 44 (7), 1255-1275.


Daniel, A., D. Birot, S. Blain, P. Tréguer, E. Menut and B. Leilde. (1995) A submersible flow-injection analyser for the in-situ determination of nitrite and nitrate in coastal waters. Marine Chemistry 51, 67-77.


Blain, S., Tréguer P. (1995) Iron(II) and iron(III) determination in seawater at the nanomolar level with on line preconcentration and spectrophotometric detection. Analytica Chimica Acta, 308, 425-432.


  Blain, S., P. Appriou and H. Handel. (1993) Preconcentration of trace metal from seawater with a chelating resin: chelamine. Analytica Chimica Acta. 272, 91-97.


Blain, S., P Appriou and H. Handel(1991). Column preconcentration of trace metals from seawater with macroporous resins impregnated with lipophilic tetraazamacrocycles. Analyst, 116, 815-820.


  Blain, S., H. Chaumeil, P. Appriou and H. Handel (1990). Application of a tetrazamacrocycle immobilized on an organic polymer to the determination of trace amounts of manganese in seawater.Analytica chimica Acta. 232, 331-336.


Invited presentation to international conferences and workshops

Blain, S. Overview of KEOPS A Synthesis of Mesoscale Iron-Enrichments, SOLAS workshop 2005 Wellington, New Zeland.

Blain, S. Natural iron fertilization in the ocean and GEOTRACES, 13-18 Jul. 2008 Goldschmidt Conference, Vancouver, Canada.

Blain, S. Nutrients availability and carbon export from the surface layer, 2008 EUROCEANS workshop, Controls on organic carbon export and twilight zone mineralization, Bruxelles, Belgique.

Blain, S. What did we learn from natural iron source? 27-28 Sept. 2011 Iron fertilization international symposium, Woods Hole, USA.

Blain, S., Impact of natural iron fertilization in the Southern Ocean, large uncertainties subsist, Chapman conference, 1-4 Sept. 2009, Southampton, UK.

Blain, S. Overview of the natural iron fertilisation experiment KEOPS, 27-29 June 2011 Symposium on Modelling and Synthesis of Southern Ocean Natural Iron Fertilization, Woods hole, USA.

Blain, S. Variability of chlorophyll, iron, and light/mixing regime in the Southern Ocean: an in situ view, May 2012, Workshop Southern Ocean overturning: stability and impacts on carbon drawdown, May 2012, Southampton, UK.

Blain S. What to consider when planning an integrated field work – modelling experiment? GEOTRACES/SCOR/SCAR Workshop, 2014 Capetown South Africa



Traductions :

    Le LOMIC en chiffres

    8 Chercheurs CNRS
    3 Enseignant-chercheurs

    1 Chercheuse accueillie

    9 Tech/Ingénieurs

    2 Post doctorants

    2 CDD Ingénieur
    12 Etudiants en thèse